Gelada follower males

Here is the abstract of brand-spanking-new research showing different reproductive strategies for adult gelada males. Geladas live in a social structure that is "harem-like", with one male mating with multiple females. However some harems have "follower males", old leader males who stuck around or younger males who didn't have what it takes to take over the unit for themselves. These potentially lower-quality follower males get some benefit, though. 18% of paternity in the unit! In return, leader males tolerate this because the presence of follower males increases leader males' tenure by 61%. This paper is presently under review.